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Abstract The history of man has always been marked by a special relationship with water. Biological function and the properties of life are often directly a result of the properties of water. It is the most abundant molecule in living organisms and it is, therefore, possible to learn a great deal about life by learning about water. On a global basis, water is a paramount factor in determining the distribution of species, and the responses and adaptation of ’species to water-stress are critical for their success in any environmental niche (Bazzaz and Fajer, 1992). Cell enlargement results from an increasing water content of the tissues. Water is absorbed by osmosis, and turgor pressure builds as the water enters. The lack of full turgor pressure keeps the water potential lower inside the cells than outside (Maruyoma and Boyer, 1994). Drought is probably the most important climatic factor limiting crop productivity world-wide and particularly in temperate regions (Jones and Corlett, 1992). Generally, water deficit reduces the growth of drought-sensitive glycophytes. Reduction in plant growth by water deficit may result from the disturbance of water balance, reduction of leaf expansion and elongation, mitotic division of cells, fresh and dry weights, changes in various physiological and metabolic processes such as decline in solute and antioxidant accumulation and expression of stress-specific genes (Hughes et aI., 1989; Munns and Sharp, 1993; Sangwan et al., 1994; Nonami et al., 1997; Schuppler et al., 1998). |