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Abstract The aim of this investigation was to determine the heterosis and types of gene action and their interaction with planting dates for some growth and yield characteristics, i.e. heading dates, plant height number of tiller and spikes per plant, height of flag region, flag leaf area, spike length, number of grains per spike, total plant weight grain yield per plant, straw yield per plant, harvest index and lOOO-kernel weight. To achieve this target F 1 of a half diallel cross between six parental genotypes namely; Sids 4 (PI), Line-l (P2), Line-2 (P3), Gemm~iza 3 (P4), Sakha 69 CPs) and Gernrneiza 1 (P6) having wide range of variability in most of the studied traits were utilized. In 1997/98 season crossing was made with all possible combinations between the six parents and hybrids and their F1’s were evaluated in successive season 1998/1999 in a randomized complete block design with three replications in three planting dates. Data were recorded on 10 guarded plants, from each plot. Analysis of. variance was performed for the studied characters in each date and then combined analysis was carried out whenever homogeneity of error variance was realized. Heterosis mean squares were significant and effects for both mid, better parent were calculated. The data were genetically analysed by the procedures of Griffing (1956), method-2, model-I. The obtained results can be summarized as follows: A- Analysis of variance, means and heterosis: 1. Planting date mean squares were significant for all the studied characters. 2. Significant genotype mean squares were detected for all studied characters in the separate planting dates as well as the combined analysis. Significant genotype x planting date interaction mean squares were obtained for all the studied characters. 3. Mean squares due to parents and hybrids were significant for all the studied characters except stem diameter in the three planting dates and in the combined analysis, lOOO-kemel weight in the third date planting date and main spike weight for parents in the second, third and the combined data and hybrids in the third planting date and lOOO-kernel weight in the first and second planting dates for hybrids. 4. Significant mean squares due to interaction between planting date and both parents and hybrids were detected for all characters except for stem diameter and lOOO-kemelweight. 5. The highest values for number of tillers and spikes per plant were detected in crosses Sakha 69 x Gemmeiza 1, Gemmeiza 3 x Gemmeiza 1, Sids 4 x Sakha 69, Gemmeiza 3 x Sakha 69. The lowest values for this trait were recorded for Line-I x Line- 2 and Sids 4 x Line-I . 6. Earlines in heading was detected in the FI hybrids of Line-l x Sids 4, Sids 4 x Line-2, Line-2 x Gemmeiza 3 and Line-2 x Gemmeiza 1. 7. Four hybrids Sids 4 x Sakha 69, Line-l x Gemmeiza 3, Sids 4 x Gemmeiza 3 and Sakha 69 x Gemmeiza 1 gave the highest mean values for grain yield per plant, number of spikes/pant, heading date, flag leaf area and lOOO-kelnel weight. Heterosis: 8. For heading date, 12 ’12, 11 and 11 eros es had significant negative heterotic effects relative to mid-parent in the first, second, third planting dates as well as the combined analysis, respectively. Significant negative heterotic effect relative to better parent was detected in crosses Gemmeiza 3 x Sakha 69 Line-I x Sakha 69 and Line-l x Gemmeiza 3. 9. For flag leaf area, 14, 15 13 and 14 hybrids exhibited significant positive heterotic effects relative to mid-parent value in the first, second and third planting dates as well as the combined analysis, respectively. 10. For number of tillers and spikes/plant, 14, 12, 14 and 14 hybrids expressed significant positive heterotic effect relative to mid-parent value in the first, second and third planting dates and the combined analysis, respectively. Also, 8, 5, 5 and 5 crosses expressed significant positive heterotic effects relative to better parent in the same order. 11. Concerning number of grains per spike tell parental combinations expressed significant positive heterotic effects over mid-parent or better parent values in the three planting dates and the combined over them. 12. For lOOO-kernel weight, aU parental combinations exhibited significant positive heterotic effect relari ve to either mid- parent or better parent values in the three planting dates and the combined analysis. 13. Regarding, grain yield per plant, 13, 11, 15 and 15 hybrids significantly exceeded mid-parent value in the first, second and third, planting dates as well as the combined analysis, respectively. Also, 10, 6, 8 and 8 hybrids from the previous parental combinations showed significant positive heterotic effects relative to better parent value in the same order. 14. For total plant weight, 13, 13 15 and 15 hybrids showed significant positive heterotic effects over the mid parent value in the first, second and third planting dates as well as the combined analysis. Also, 8, 8, 11 and 9 crosses from the previous parental combinations gave significant positive heterotic effects relative to better parent value in the same order. 15. For harvest index, 3, 8, 7 and 6 hybrids showed significant positive heterotic effects relative to mid-parent in the first second. and third planting dates as well as the combined analysis. Also, 3, 4, 5 and 4 hybrids from the previous crosse’; expressed significantly positive heterotic effects over better parent in the same order. B- Combining ability: 16. The mean squares associated with general combining ability (G.C.A.) and specific combining ability (S.C.A.) were significant for all studied characters. Low G. .A./S.c.A. ratios of less than uni ty were detected for stem diameter for the three planting dates as well as for combined analysis. Moreover, for lOOO-kernel weight in the se and and third planting dates, weight of main spike in the third planting date and harvest index in the fast, second planting date and the combined data they were also significant. Low G.C.A./S.C.A ratio of less than unity indicated the functioning of non-adrutive gene action in the inheritance of such characters. On the other hand, high G.C.A.lS.C.A. ratio which exceeded the unity were detected for other cases. Such results indicated that additive and additive x additive types of gene actions which are more important than non-additive gene actions in controlling these characters. 17. The mean squares of interaction between planting dates and both types of combining ability were significant for all traits except stem diameter, revealing that the magnitude of additive, additive x additive and non-additive types of gene action varied from planting dates to another. General combining ability effects: 18. Line-2 could be considered as an excellent parent in breeding programs towards releasing varieties with higher of plant height, stem diameter and flag leaf area. 19. Parental varieties Sids 4, Line-I and Line-2 expressed significant negative effects for heading date in the three planting dates as well as the the combined analysis, indicating that the three varieties could be considered as good combiners for developing breading programe for earliness. 20. For number of spikes and tillers/plant Sakha 69 and Gemmeiza 1 could be considered as an excellent parents in breeding programes towards releasing varieties characterized by high number of spikes/plant. 21. Sids 4 seemed to be good combiner for number of kemeJ.s/spike and spike length. 22. For grain and straw yields and total plant weight Sakha 69 gave the highest significant positive effects followed by Gemmeiza 1 and then by Gemmeiza 3 in the three planting dates as well as in the combined data for these character and some of their components. 23. Significant correlation coefficient values between the parental performance and its (gi) effects were obtained for heading date, plant eight, number of spikes and tillers per plant, spike length, main spike weight, straw and grain yield/plant in the three planting dates as well as the combined analysis, nurnber of grains/spike ill the third planting date and in the combined analysis. Likeness, the same result was obtained for peduncle length in the second, third planting dates and the combined analysis and stem diameter in the first, second planting date and the combined analysis. Specific combining ability effects: 24. The crosses Line-2 x Gemmeiza 1. Sids 4 x Gemmeiza 1 and Line-I x Sakha 69 gave the highest desirable CSij) effects for heading ate in the three planting dates and in the combined analysis. 25. Crosses Sids 4 x Line-2, Sids 4 x Gemmeiza 3, Sids 4 x Sakha 69, Sids 4 x Gemmeiza 1, Line-I x Gemmeiza 3, Line-2 x Gemmeiza 3, Line-2 x Gernmeiza 1 gave significant positive CSiJ effects for grain, straw yields/plant and total plant weight in the three planting dates and in the combined analysis. 26. The cross Sids 4 x Line-l had the highest positive value (Sij) effect followed by Genuneiza 3 x Saka-69 and then by cross Line-1 x Line-2 in the combined analysis for flag leaf area. 27. Regarding plant height, eight eleven, eleven and twelve crosses exhibited significant positive CCSij) effects in the three planting dates and in the combined analysis. The highest positive effects was obtained from crosses Line-2 x Gemmeiza 1, Sids 4 x Line-I, Line-2 x Sakha 69 and Line- 1 x Gemmeiza 3 in the combined analysis. However 2, 2, 3 and 3 crosses expressed significant negative effects in the same order. The best cross was Gemmeiza 3 x Salcha 69 which gave the lowest value of (5;) effect in the combined analysis. 28. For number of tillers and spikes per plant, the highest positive (Sij) effects was detected for the three crosses Sids 4 x Gemmeiza 1, Line-2 x Gemmeiza 1 and Sids 4 x Gemmeiza 3 in the combined analysis for both traits. 29. For peduncle length, the best combinations were Line-l x Gemmeiza 3, Line-2 x Gemmeiza 1 and Sakha 69 x Gemmeiza 1 in the combined analysis. 30. For spike length and number of grains/spike the best crosses were Line-2 x Genuneiza 1 and Sakha 69 x Gemmeiza 1 and had the highest positive value of (Sij) effect in the combined analysis for both traits. 31. The two crosses Line-I x Line-2 and l.ine-I x Gemmeiza 3 showed the highest positive val ues of (Sjj) in the combined analysis for weight of main spike. 32. For harvest index, the three crosses Sids 4 x Line-I, Gemmeiza 1 x Gemrneiza 3 and Gemmeiza 3 x Sakha 69, gave the highest (Sij) values in the combined analysis. |